We reveal a near-complete 70-million-year-old crocodile relative from Argentina—Kostensuchus atrox —and what its broad snout, giant jaw muscles, and serrated teeth tell us about Cretaceous land predation. The episode compares its ecology with coeval predators like Chaetotrox and Baryosuchids, explores locomotor clues from its limbs, and explains why this extraordinary fossil reshapes our view of top predators on the eve of the dinosaur extinction.

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