Long before T. rex, croc-line archosaurs—the pseudosuchians—dominated the Triassic. We trace their game-changing crurotarsal ankle that allowed both sprawling locomotion and an erect, powerful stance, explore iconic predators like Postosuchus and Desmatosuchus, and spotlight the Brazil-born Papasaurid Tenraclusucus bellator as a case study in early reptile innovation. Finally, we connect these fossils to the bigger story of life's reshaping end-Triassic extinction and what they reveal about the deep, surprising history of dominance and discovery.

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