In a settlement known as Stonewall, two bunnies are given a mission by Grandma Maizy, the matriarch. Jon Radisher, a radish trader, has disappeared along the route to another burrow known as Bywater. Foo Foo, a fighter, and Run E. Carlos, a runner, begin their search for the missing bunny immediately, soon coming upon their first clue as well as a mischievous gang of weasels.