Koko is perhaps the world's most famous gorilla. She was known for her ability to communicate using thousands of signs and gestures taught to her by her caretaker, Francine "Penny" Patterson as part of a research project that spanned 40 years. KoKo captured the public imagination with the way she appeared to think and emote like a human. But as Koko’s fame grew, animal behavioral theorists started to have doubts about the study’s method. How much did Koko really know? What did Koko the talking gorilla really have to say?

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