Alex Kingston's career really took off in the mid-1990s, first portraying Dr. Elizabeth Corday on NBC's ER, and then in the late 2000s, as River Song, the mysterious friend who knew Doctor Who's future on the hit BBC series. Spoilers! All of it followed a devastating split from her partner of 12 years, and husband of two years, Ralph Fiennes. While playing Hamlet, Fiennes fell in love with the actress playing Hamlet's mother Gertrude - launching a decade long relationship that would itself implode amid infidelity allegations.

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