Frans de Waal is a Dutch-American primatologist and the Charles Howard Candler Professor of Primate Behavior in the Department of Psychology at Emory University. He is the author of, among other books, Chimpanzee Politics: Power and Sex Among Apes and Different: Gender Through the Eyes of a Primatologist.

In this week’s conversation, Yascha Mounk and Frans de Waal discuss how primate empathy forms the basis for human morality; the precedent for diversity in human gender and gender roles that exists in non-human primates; and why, despite attempts to separate human behavior from human biology, you can’t have one without the other.

This transcript has been condensed and lightly edited for clarity.

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