After the shocking attack on Sardis, many more Greek cities joined the Ionian Revolt, despite Persian victory at Ephesus. In 497 BCE, three land campaigns were launched by three Persian generals: Daurises, Hymaies, and Otanes. After a series of lightning victories in early 497, the campaigns began set in to prolonged fighting. Two of the Persian generals were dead by 496, but the Ionians were still losing. Fresh revolts in the Troad and Caria were dealt serious defeats, and Aristagoras of Miletus, once the ringleader of the Ionians, fled into exile.

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