Richard Coles and Anita Anand with neuroscientist Baroness Susan Greenfield talking about her life and work, Julie Bailey who campaigned for reform after her Mother died at Stafford Hospital and who has been hounded from her Stafford home and business, 'Lorna' who lives in Belfast, was in care from around 14 years of age and now volunteers with a Children in Need project that helps others in care, and Diccon Bewes who has followed in the diary footsteps of Miss Jemima Morrell from Selby in Yorkshire, who took an 1863 tour of Switzerland organised by Thomas Cook. JP Devlin visits the village of Herbrandston in Wales which is a doubly 'Thankful Village'- all the village soldiers returned from both World Wars. Back in the studio, Medwyn Parry talks about his motorbike ride to 51 Thankful Villages in the UK- and tells of two new ones discovered recently. And Gary Barlow shares his Inheritance Tracks.

Producer: Chris Wilson.